JOINT FILING AGREEMENT

Exhibit 99.1

JOINT FILING AGREEMENT

We, the undersigned, hereby express our agreement that the attached Schedule 13G is filed on behalf of each of us.

Dated: February 11, 2019

THE COLUMN GROUP, LP				THE COLUMN GROUP GP, LP
By:		The Column Group GP, LP
By:		/s/ Xxxxx Xxxxxxxxxx		By:		/s/ Xxxxx Xxxxxxxxxx
Name:		Xxxxx Xxxxxxxxxx		Name:		Xxxxx Xxxxxxxxxx
Title:		Managing Partner		Title:		Managing Partner
PONOI CAPITAL, LP				PONOI MANAGEMENT, LLC
By:		Ponoi Management, LLC
By:		/s/ Xxxxx Xxxxxxxxxx		By:		/s/ Xxxxx Xxxxxxxxxx
Name:		Xxxxx Xxxxxxxxxx		Name:		Xxxxx Xxxxxxxxxx
Title:		Managing Partner		Title:		Managing Partner
PONOI CAPITAL II, LP				PONOI II MANAGEMENT, LLC
By:		Ponoi II Management, LLC
By:		/s/ Xxxxx Xxxxxxxxxx		By:		/s/ Xxxxx Xxxxxxxxxx
Name:		Xxxxx Xxxxxxxxxx		Name:		Xxxxx Xxxxxxxxxx
Title:		Managing Partner		Title:		Managing Partner
By:		/s/ Xxxxx Xxxxxxxxxx
		Xxxxx Xxxxxxxxxx
By:		/s/ Xxxxx X. Xxxxxxx
		Xxxxx X. Xxxxxxx
By:		/s/ Xxx Xxxxxxx
		Xxx Xxxxxxx