CELLECTIS S.A. (a société anonyme incorporated in the French Republic) 5,646,000 American Depositary Shares Each Representing One Ordinary Share (nominal value €0.05 per share) UNDERWRITING AGREEMENT Dated: April 4, 2018Underwriting Agreement • April 10th, 2018 • Cellectis S.A. • Biological products, (no disgnostic substances) • New York
